Evaluating models of climate and forest vegetation

Understanding how the biosphere may respond to increasing trace gas concentrations in the atmosphere requires models that contain vegetation responses to regional climate. Most of the processes ecologists study in forests, including trophic interactions, nutrient cycling, and disturbance regimes, and vital components of the world economy, such as forest products and agriculture, will be influenced in potentially unexpected ways by changing climate. These vegetation changes affect climate in the following ways: changing C, N, and S pools; trace gases; albedo; and water balance. The complexity of the indirect interactions among variables that depend on climate, together with the range of different space/time scales that best describe these processes, make the problems of modeling and prediction enormously difficult. These problems of predicting vegetation response to climate warming and potential ways of testing model predictions are the subjects of this chapter.